The problem: In the real world, GNSS isn't always available.
Jamming, spoofing, tunnels, urban canyon, indoors, underwater. Anywhere satellite positioning stops being trustworthy.
When the fix goes, the platform still has to know how it is moving. The MK2E2 and MK2Z pair an optical gyro axis with a full MEMS inertial suite. The MK2M2 runs that MEMS suite alone where size and cost lead.